Ichthyology conducts research, research training and graduate education on the world’s fishes – their global diversity, evolution, geography, genomics, morphology, conservation, ecology and behavior. These studies are grounded in research collections of 660,000 preserved and osteological specimens and 9,500 tissue samples, which provide a major world resource for genomic and molecular systematics of fishes. Curator Ed Wiley focuses on higher level evolutionary relationships of fishes and the theory and practice of phylogenetic systematics. Other researchers in Ichthyology study the evolution and diversity of fossil and living fishes.